List of experimentally validated KVxF/RVxF motif instances curated from the PP1 literature. All instances have been annotated with their publication and stuctural information if available. All data is clickable to access source data.

References

(1) "The anti-apoptotic molecules Bcl-xL and Bcl-w target protein phosphatase 1alpha to Bad." Ayllón V, Cayla X, García A, Fleischer A, Rebollo A. Eur. J. Immunol.. 2002 Jul;32 (7):1847-55

(2) "Multiple interactions within the AKAP220 signaling complex contribute to protein phosphatase 1 regulation." Schillace RV, Voltz JW, Sim AT, Shenolikar S, Scott JD. J. Biol. Chem.. 2001 Apr;276 (15):12128-34

(3) "Characterization of a novel giant scaffolding protein, CG-NAP, that anchors multiple signaling enzymes to centrosome and the golgi apparatus." Takahashi M, Shibata H, Shimakawa M, Miyamoto M, Mukai H, Ono Y. J. Biol. Chem.. 1999 Jun;274 (24):17267-74

(4) "Structural analysis of the protein phosphatase 1 docking motif: molecular description of binding specificities identifies interacting proteins." Meiselbach H, Sticht H, Enz R. Chem. Biol.. 2006 Jan;13 (1):49-59

(5) "SARP, a new alternatively spliced protein phosphatase 1 and DNA interacting protein." Browne GJ, Fardilha M, Oxenham SK, Wu W, Helps NR, da Cruz E Silva OA, Cohen PT, da Cruz E Silva EF. Biochem. J.. 2007 Feb;402 (1):187-96

(6) "Bcl-2 targets protein phosphatase 1 alpha to Bad." Ayllón V, Cayla X, García A, Roncal F, Fernández R, Albar JP, Martínez C, Rebollo A. J. Immunol.. 2001 Jun;166 (12):7345-52

(7) "Protein phosphatase 1 interacts with p53BP2, a protein which binds to the tumour suppressor p53." Helps NR, Barker HM, Elledge SJ, Cohen PT. FEBS Lett.. 1995 Dec;377 (3):295-300

(8) "Interaction and feedback regulation between STK15/BTAK/Aurora-A kinase and protein phosphatase 1 through mitotic cell division cycle." Katayama H, Zhou H, Li Q, Tatsuka M, Sen S. J. Biol. Chem.. 2001 Dec;276 (49):46219-24

(9) "Aurora-B associated protein phosphatases as negative regulators of kinase activation." Sugiyama K, Sugiura K, Hara T, Sugimoto K, Shima H, Honda K, Furukawa K, Yamashita S, Urano T. Oncogene. 2002 May;21 (20):3103-11

(10) "Identification and functional characterization of a PP1-binding site in BRCA1." , , , , . Biochem. Biophys. Res. Commun.. 2007 Aug;360 (2):507-12

(11) "Repo-Man recruits PP1 gamma to chromatin and is essential for cell viability." Trinkle-Mulcahy L, Andersen J, Lam YW, Moorhead G, Mann M, Lamond AI. J. Cell Biol.. 2006 Feb;172 (5):679-92

(12) "The Ki-67 and RepoMan mitotic phosphatases assemble via an identical, yet novel mechanism." Kumar GS, Gokhan E, De Munter S, Bollen M, Vagnarelli P, Peti W, Page R. Elife. 2016 08;5 ():

(13) "Aurora kinases and protein phosphatase 1 mediate chromosome congression through regulation of CENP-E." Kim Y, Holland AJ, Lan W, Cleveland DW. Cell. 2010 Aug;142 (3):444-55

(14) "Aurora B opposes PP1 function in mitosis by phosphorylating the conserved PP1-binding RVxF motif in PP1 regulatory proteins." Nasa I, Rusin SF, Kettenbach AN, Moorhead GB. Sci Signal. 2018 May;11 (530):

(15) "The RNA-binding protein Staufen from rat brain interacts with protein phosphatase-1." Monshausen M, Rehbein M, Richter D, Kindler S. J. Neurochem.. 2002 May;81 (3):557-64

(16) "Identification and characterization of the human HCG V gene product as a novel inhibitor of protein phosphatase-1." Zhang J, Zhang L, Zhao S, Lee EY. Biochemistry. 1998 Nov;37 (47):16728-34

(17) "IPP5, a novel protein inhibitor of protein phosphatase 1, promotes G1/S progression in a Thr-40-dependent manner." Wang X, Liu B, Li N, Li H, Qiu J, Zhang Y, Cao X. J. Biol. Chem.. 2008 May;283 (18):12076-84

(18) "FLJ23654 encodes a heart protein phosphatase 1-binding protein (Hepp1)." Chen CY, Lai NS, Yang JJ, Huang HL, Hung WC, Li C, Lin TH, Huang HB. Biochem. Biophys. Res. Commun.. 2010 Jan;391 (1):698-702

(19) "The translation initiation factor eIF2beta is an interactor of protein phosphatase-1." Wakula P, Beullens M, van Eynde A, Ceulemans H, Stalmans W, Bollen M. Biochem. J.. 2006 Dec;400 (2):377-83

(20) "Interaction of p130 with, and consequent inhibition of, the catalytic subunit of protein phosphatase 1alpha." Yoshimura K, Takeuchi H, Sato O, Hidaka K, Doira N, Terunuma M, Harada K, Ogawa Y, Ito Y, Kanematsu T, Hirata M. J. Biol. Chem.. 2001 May;276 (21):17908-13

(21) "The human mitotic kinesin KIF18A binds protein phosphatase 1 (PP1) through a highly conserved docking motif." De Wever V, Nasa I, Chamousset D, Lloyd D, Nimick M, Xu H, Trinkle-Mulcahy L, Moorhead GB. Biochem. Biophys. Res. Commun.. 2014 Oct;453 (3):432-7

(22) "PP1-Mediated Dephosphorylation of Lgl Controls Apical-basal Polarity." Moreira S, Osswald M, Ventura G, Gonçalves M, Sunkel CE, Morais-de-Sá E. Cell Rep. 2019 Jan;26 (2):293-301.e7

(23) "Group I metabotropic glutamate receptors bind to protein phosphatase 1C. Mapping and modeling of interacting sequences." Croci C, Sticht H, Brandstätter JH, Enz R. J. Biol. Chem.. 2003 Dec;278 (50):50682-90

(24) "Development of a peptide that selectively activates protein phosphatase-1 in living cells." Chatterjee J, Beullens M, Sukackaite R, Qian J, Lesage B, Hart DJ, Bollen M, Köhn M. Angew. Chem. Int. Ed. Engl.. 2012 Oct;51 (40):10054-9

(25) "Regulation of neurabin I interaction with protein phosphatase 1 by phosphorylation." McAvoy T, Allen PB, Obaishi H, Nakanishi H, Takai Y, Greengard P, Nairn AC, Hemmings HC. Biochemistry. 1999 Sep;38 (39):12943-9

(26) "Brain actin-associated protein phosphatase 1 holoenzymes containing spinophilin, neurabin, and selected catalytic subunit isoforms." MacMillan LB, Bass MA, Cheng N, Howard EF, Tamura M, Strack S, Wadzinski BE, Colbran RJ. J. Biol. Chem.. 1999 Dec;274 (50):35845-54

(27) "Spinophilin directs protein phosphatase 1 specificity by blocking substrate binding sites." Ragusa MJ, Dancheck B, Critton DA, Nairn AC, Page R, Peti W. Nat. Struct. Mol. Biol.. 2010 Apr;17 (4):459-64

(28) "" . . ; ():

(29) "Spinophilin, a novel protein phosphatase 1 binding protein localized to dendritic spines." Allen PB, Ouimet CC, Greengard P. Proc. Natl. Acad. Sci. U.S.A.. 1997 Sep;94 (18):9956-61

(30) "Site-specific dephosphorylation of doublecortin (DCX) by protein phosphatase 1 (PP1)." Shmueli A, Gdalyahu A, Sapoznik S, Sapir T, Tsukada M, Reiner O. Mol. Cell. Neurosci.. ;32 (1-2):15-26

(31) "Val193 and Phe195 of the gamma 1 34.5 protein of herpes simplex virus 1 are required for viral resistance to interferon-alpha/beta." Cheng G, Brett ME, He B. Virology. 2001 Nov;290 (1):115-20

(32) "Consensus PP1 binding motifs regulate transcriptional corepression and alternative RNA splicing activities of the steroid receptor coregulators, p54nrb and PSF." Liu L, Xie N, Rennie P, Challis JR, Gleave M, Lye SJ, Dong X. Mol. Endocrinol.. 2011 Jul;25 (7):1197-210

(32) "Consensus PP1 binding motifs regulate transcriptional corepression and alternative RNA splicing activities of the steroid receptor coregulators, p54nrb and PSF." Liu L, Xie N, Rennie P, Challis JR, Gleave M, Lye SJ, Dong X. Mol. Endocrinol.. 2011 Jul;25 (7):1197-210

(33) "Functional interaction between nuclear inhibitor of protein phosphatase type 1 (NIPP1) and protein phosphatase type 1 (PP1) in Drosophila: consequences of over-expression of NIPP1 in flies and suppression by co-expression of PP1." Parker L, Gross S, Beullens M, Bollen M, Bennett D, Alphey L. Biochem. J.. 2002 Dec;368 (Pt 3):789-97

(34) "Molecular determinants of nuclear protein phosphatase-1 regulation by NIPP-1." Beullens M, Van Eynde A, Vulsteke V, Connor J, Shenolikar S, Stalmans W, Bollen M. J. Biol. Chem.. 1999 May;274 (20):14053-61

(35) "The molecular basis for substrate specificity of the nuclear NIPP1:PP1 holoenzyme." O'Connell N, Nichols SR, Heroes E, Beullens M, Bollen M, Peti W, Page R. Structure. 2012 Oct;20 (10):1746-56

(36) "N-CoR controls differentiation of neural stem cells into astrocytes." Hermanson O, Jepsen K, Rosenfeld MG. Nature. 2002 Oct;419 (6910):934-9

(37) "Association of a protein phosphatase 1 activity with the human factor C1 (HCF) complex." Ajuh PM, Browne GJ, Hawkes NA, Cohen PT, Roberts SG, Lamond AI. Nucleic Acids Res.. 2000 Feb;28 (3):678-86

(38) "NOM1 targets protein phosphatase I to the nucleolus." Gunawardena SR, Ruis BL, Meyer JA, Kapoor M, Conklin KF. J. Biol. Chem.. 2008 Jan;283 (1):398-404

(39) "RSV hijacks cellular protein phosphatase 1 to regulate M2-1 phosphorylation and viral transcription." Richard CA, Rincheval V, Lassoued S, Fix J, Cardone C, Esneau C, Nekhai S, Galloux M, Rameix-Welti MA, Sizun C, Eléouët JF. PLoS Pathog.. 2018 03;14 (3):e1006920

(40) "Identification of phostensin, a PP1 F-actin cytoskeleton targeting subunit." Kao SC, Chen CY, Wang SL, Yang JJ, Hung WC, Chen YC, Lai NS, Liu HT, Huang HL, Chen HC, Lin TH, Huang HB. Biochem. Biophys. Res. Commun.. 2007 May;356 (3):594-8

(41) "Channel-anchored protein kinase CK2 and protein phosphatase 1 reciprocally regulate KCNQ2-containing M-channels via phosphorylation of calmodulin." Kang S, Xu M, Cooper EC, Hoshi N. J. Biol. Chem.. 2014 Apr;289 (16):11536-44

(42) "Ki-67 is a PP1-interacting protein that organises the mitotic chromosome periphery." Booth DG, Takagi M, Sanchez-Pulido L, Petfalski E, Vargiu G, Samejima K, Imamoto N, Ponting CP, Tollervey D, Earnshaw WC, Vagnarelli P. Elife. 2014 May;3 ():e01641

(43) "A Nucleoporin Docks Protein Phosphatase 1 to Direct Meiotic Chromosome Segregation and Nuclear Assembly." Hattersley N, Cheerambathur D, Moyle M, Stefanutti M, Richardson A, Lee KY, Dumont J, Oegema K, Desai A. Dev. Cell. 2016 09;38 (5):463-77

(44) "Phosphorylation of TIMAP by glycogen synthase kinase-3beta activates its associated protein phosphatase 1." Li L, Kozlowski K, Wegner B, Rashid T, Yeung T, Holmes C, Ballermann BJ. J. Biol. Chem.. 2007 Aug;282 (35):25960-9

(45) "Structural basis of protein phosphatase 1 regulation." Terrak M, Kerff F, Langsetmo K, Tao T, Dominguez R. Nature. 2004 Jun;429 (6993):780-4

(46) "AKAP149 is a novel PP1 specifier required to maintain nuclear envelope integrity in G1 phase." Steen RL, Beullens M, Landsverk HB, Bollen M, Collas P. J. Cell. Sci.. 2003 Jun;116 (Pt 11):2237-46

(47) "The major myosin phosphatase in skeletal muscle is a complex between the beta-isoform of protein phosphatase 1 and the MYPT2 gene product." Moorhead G, Johnson D, Morrice N, Cohen P. FEBS Lett.. 1998 Nov;438 (3):141-4

(48) "Phosphorylation of a novel myosin binding subunit of protein phosphatase 1 reveals a conserved mechanism in the regulation of actin cytoskeleton." Tan I, Ng CH, Lim L, Leung T. J. Biol. Chem.. 2001 Jun;276 (24):21209-16

(49) "GBPI, a novel gastrointestinal- and brain-specific PP1-inhibitory protein, is activated by PKC and inactivated by PKA." Liu QR, Zhang PW, Lin Z, Li QF, Woods AS, Troncoso J, Uhl GR. Biochem. J.. 2004 Jan;377 (Pt 1):171-81

(50) "Feedback inhibition of the unfolded protein response by GADD34-mediated dephosphorylation of eIF2alpha." Novoa I, Zeng H, Harding HP, Ron D. J. Cell Biol.. 2001 May;153 (5):1011-22

(51) "Structural and Functional Analysis of the GADD34:PP1 eIF2α Phosphatase." Choy MS, Yusoff P, Lee IC, Newton JC, Goh CW, Page R, Shenolikar S, Peti W. Cell Rep. 2015 Jun;11 (12):1885-91

(52) "G-actin provides substrate-specificity to eukaryotic initiation factor 2α holophosphatases." Chen R, Rato C, Yan Y, Crespillo-Casado A, Clarke HJ, Harding HP, Marciniak SJ, Read RJ, Ron D. Elife. 2015 Mar;4 ():

(53) "Cloning and identification of MYPT3: a prenylatable myosin targetting subunit of protein phosphatase 1." Skinner JA, Saltiel AR. Biochem. J.. 2001 May;356 (Pt 1):257-67

(54) "Multiple structural elements define the specificity of recombinant human inhibitor-1 as a protein phosphatase-1 inhibitor." Endo S, Zhou X, Connor J, Wang B, Shenolikar S. Biochemistry. 1996 Apr;35 (16):5220-8

(55) "DARPP-32, a dopamine-regulated neuronal phosphoprotein, is a potent inhibitor of protein phosphatase-1." Hemmings HC, Greengard P, Tung HY, Cohen P. Nature. ;310 (5977):503-5

(56) "Protein phosphatase-1 binding to scd5p is important for regulation of actin organization and endocytosis in yeast." Chang JS, Henry K, Wolf BL, Geli M, Lemmon SK. J. Biol. Chem.. 2002 Dec;277 (50):48002-8

(57) "Identification of protein-phosphatase-1-binding domains on the glycogen and myofibrillar targetting subunits." Johnson DF, Moorhead G, Caudwell FB, Cohen P, Chen YH, Chen MX, Cohen PT. Eur. J. Biochem.. 1996 Jul;239 (2):317-25

(58) "Amino acid sequence and expression of the hepatic glycogen-binding (GL)-subunit of protein phosphatase-1." Doherty MJ, Moorhead G, Morrice N, Cohen P, Cohen PT. FEBS Lett.. 1995 Nov;375 (3):294-8

(59) "Role of protein targeting to glycogen (PTG) in the regulation of protein phosphatase-1 activity." Brady MJ, Printen JA, Mastick CC, Saltiel AR. J. Biol. Chem.. 1997 Aug;272 (32):20198-204

(60) "PPP1R6, a novel member of the family of glycogen-targetting subunits of protein phosphatase 1." Armstrong CG, Browne GJ, Cohen P, Cohen PT. FEBS Lett.. 1997 Nov;418 (1-2):210-4

(61) "Binding of the concave surface of the Sds22 superhelix to the alpha 4/alpha 5/alpha 6-triangle of protein phosphatase-1." Ceulemans H, Vulsteke V, De Maeyer M, Tatchell K, Stalmans W, Bollen M. J. Biol. Chem.. 2002 Dec;277 (49):47331-7

(62) "Identification of a Plasmodium falciparum inhibitor-2 motif involved in the binding and regulation activity of protein phosphatase type 1." Fréville A, Tellier G, Vandomme A, Pierrot C, Vicogne J, Cantrelle FX, Martoriati A, Cailliau-Maggio K, Khalife J, Landrieu I. FEBS J.. 2014 Oct;281 (19):4519-34

(63) "Identification and characterization of AtI-2, an Arabidopsis homologue of an ancient protein phosphatase 1 (PP1) regulatory subunit." Templeton GW, Nimick M, Morrice N, Campbell D, Goudreault M, Gingras AC, Takemiya A, Shimazaki K, Moorhead GB. Biochem. J.. 2011 Apr;435 (1):73-83

(63) "Identification and characterization of AtI-2, an Arabidopsis homologue of an ancient protein phosphatase 1 (PP1) regulatory subunit." Templeton GW, Nimick M, Morrice N, Campbell D, Goudreault M, Gingras AC, Takemiya A, Shimazaki K, Moorhead GB. Biochem. J.. 2011 Apr;435 (1):73-83

(64) "Structural basis for regulation of protein phosphatase 1 by inhibitor-2." Hurley TD, Yang J, Zhang L, Goodwin KD, Zou Q, Cortese M, Dunker AK, DePaoli-Roach AA. J. Biol. Chem.. 2007 Sep;282 (39):28874-83

(65) "BH-protocadherin-c, a member of the cadherin superfamily, interacts with protein phosphatase 1 alpha through its intracellular domain." Yoshida K, Watanabe M, Kato H, Dutta A, Sugano S. FEBS Lett.. 1999 Oct;460 (1):93-8

(66) "Functional analysis of the yeast Glc7-binding protein Reg1 identifies a protein phosphatase type 1-binding motif as essential for repression of ADH2 expression." Dombek KM, Voronkova V, Raney A, Young ET. Mol. Cell. Biol.. 1999 Sep;19 (9):6029-40

(67) "PP1 phosphatase-binding motif in Reg1 protein of Saccharomyces cerevisiae is required for interaction with both the PP1 phosphatase Glc7 and the Snf1 protein kinase." Tabba S, Mangat S, McCartney R, Schmidt MC. Cell. Signal.. 2010 Jul;22 (7):1013-21

(68) "An overlapping kinase and phosphatase docking site regulates activity of the retinoblastoma protein." Hirschi A, Cecchini M, Steinhardt RC, Schamber MR, Dick FA, Rubin SM. Nat. Struct. Mol. Biol.. 2010 Sep;17 (9):1051-7

(69) "RRP1B targets PP1 to mammalian cell nucleoli and is associated with Pre-60S ribosomal subunits." Chamousset D, De Wever V, Moorhead GB, Chen Y, Boisvert FM, Lamond AI, Trinkle-Mulcahy L. Mol. Biol. Cell. 2010 Dec;21 (23):4212-26

(70) "Binding of the catalytic subunit of protein phosphatase-1 to the ryanodine-sensitive calcium release channel protein." Zhao S, Brandt NR, Caswell AH, Lee EY. Biochemistry. 1998 Dec;37 (51):18102-9

(71) "Protein phosphatase-1 is a novel regulator of the interaction between IRBIT and the inositol 1,4,5-trisphosphate receptor." Devogelaere B, Beullens M, Sammels E, Derua R, Waelkens E, van Lint J, Parys JB, Missiaen L, Bollen M, De Smedt H. Biochem. J.. 2007 Oct;407 (2):303-11

(72) "A Phytophthora infestans RXLR effector targets plant PP1c isoforms that promote late blight disease." Boevink PC, Wang X, McLellan H, He Q, Naqvi S, Armstrong MR, Zhang W, Hein I, Gilroy EM, Tian Z, Birch PR. Nat Commun. 2016 Jan;7 ():10311

(73) "Hipk2 and PP1c cooperate to maintain Dvl protein levels required for Wnt signal transduction." Shimizu N, Ishitani S, Sato A, Shibuya H, Ishitani T. Cell Rep. 2014 Sep;8 (5):1391-404

(74) "Apoptosis-associated tyrosine kinase scaffolding of protein phosphatase 1 and SPAK reveals a novel pathway for Na-K-2C1 cotransporter regulation." Gagnon KB, England R, Diehl L, Delpire E. Am. J. Physiol., Cell Physiol.. 2007 May;292 (5):C1809-15

(75) "NIMA-related kinase 2 (Nek2), a cell-cycle-regulated protein kinase localized to centrosomes, is complexed to protein phosphatase 1." Helps NR, Luo X, Barker HM, Cohen PT. Biochem. J.. 2000 Jul;349 (Pt 2):509-18

(76) "PNUTS, a protein phosphatase 1 (PP1) nuclear targeting subunit. Characterization of its PP1- and RNA-binding domains and regulation by phosphorylation." Kim YM, Watanabe T, Allen PB, Kim YM, Lee SJ, Greengard P, Nairn AC, Kwon YG. J. Biol. Chem.. 2003 Apr;278 (16):13819-28

(77) "Understanding the antagonism of retinoblastoma protein dephosphorylation by PNUTS provides insights into the PP1 regulatory code." Choy MS, Hieke M, Kumar GS, Lewis GR, Gonzalez-DeWhitt KR, Kessler RP, Stein BJ, Hessenberger M, Nairn AC, Peti W, Page R. Proc. Natl. Acad. Sci. U.S.A.. 2014 Mar;111 (11):4097-102

(78) "Purification and characterisation of p99, a nuclear modulator of protein phosphatase 1 activity." Kreivi JP, Trinkle-Mulcahy L, Lyon CE, Morrice NA, Cohen P, Lamond AI. FEBS Lett.. 1997 Dec;420 (1):57-62

(79) "Characterization of the inhibition of protein phosphatase-1 by DARPP-32 and inhibitor-2." Huang HB, Horiuchi A, Watanabe T, Shih SR, Tsay HJ, Li HC, Greengard P, Nairn AC. J. Biol. Chem.. 1999 Mar;274 (12):7870-8

(80) "PITK, a PP1 targeting subunit that modulates the phosphorylation of the transcriptional regulator hnRNP K." Kwiek NC, Thacker DF, Datto MB, Megosh HB, Haystead TA. Cell. Signal.. 2006 Oct;18 (10):1769-78

(81) "Modulation of ion transport by direct targeting of protein phosphatase type 1 to the Na-K-Cl cotransporter." Darman RB, Flemmer A, Forbush B. J. Biol. Chem.. 2001 Sep;276 (37):34359-62

(82) "Interaction of protein phosphatase type 1 with a splicing factor." Hirano K, Erdödi F, Patton JG, Hartshorne DJ. FEBS Lett.. 1996 Jul;389 (2):191-4

(83) "Taperin (c9orf75), a mutated gene in nonsyndromic deafness, encodes a vertebrate specific, nuclear localized protein phosphatase one alpha (PP1α) docking protein." Ferrar T, Chamousset D, De Wever V, Nimick M, Andersen J, Trinkle-Mulcahy L, Moorhead GB. Biol Open. 2012 Feb;1 (2):128-39

(84) "Rif1 controls DNA replication timing in yeast through the PP1 phosphatase Glc7." Mattarocci S, Shyian M, Lemmens L, Damay P, Altintas DM, Shi T, Bartholomew CR, Thomä NH, Hardy CF, Shore D. Cell Rep. 2014 Apr;7 (1):62-9

(85) "Human RIF1 and protein phosphatase 1 stimulate DNA replication origin licensing but suppress origin activation." Hiraga SI, Ly T, Garzón J, Hořejší Z, Ohkubo YN, Endo A, Obuse C, Boulton SJ, Lamond AI, Donaldson AD. EMBO Rep.. 2017 03;18 (3):403-419

(86) "Mouse Rif1 is a regulatory subunit of protein phosphatase 1 (PP1)." Sukackaite R, Cornacchia D, Jensen MR, Mas PJ, Blackledge M, Enervald E, Duan G, Auchynnikava T, Köhn M, Hart DJ, Buonomo SBC. Sci Rep. 2017 05;7 (1):2119

(87) "Tensin1 requires protein phosphatase-1alpha in addition to RhoGAP DLC-1 to control cell polarization, migration, and invasion." Hall EH, Daugherty AE, Choi CK, Horwitz AF, Brautigan DL. J. Biol. Chem.. 2009 Dec;284 (50):34713-22

(88) "PPP1CC2 can form a kinase/phosphatase complex with the testis-specific proteins TSSK1 and TSKS in the mouse testis." MacLeod G, Shang P, Booth GT, Mastropaolo LA, Manafpoursakha N, Vogl AW, Varmuza S. Reproduction. 2014 Jan;147 (1):1-12

(89) "TMEM225: a possible protein phosphatase 1γ2 (PP1γ2) regulator localizes to the equatorial segment in mouse spermatozoa." Matsuura M, Yogo K. Mol. Reprod. Dev.. 2015 Feb;82 (2):139-48

(90) "Downregulation of Fer induces PP1 activation and cell-cycle arrest in malignant cells." Pasder O, Shpungin S, Salem Y, Makovsky A, Vilchick S, Michaeli S, Malovani H, Nir U. Oncogene. 2006 Jul;25 (30):4194-206

(91) "SIPP1, a novel pre-mRNA splicing factor and interactor of protein phosphatase-1." Llorian M, Beullens M, Andrés I, Ortiz JM, Bollen M. Biochem. J.. 2004 Feb;378 (Pt 1):229-38

(92) "The nuclear PP1 interacting protein ZAP3 (ZAP) is a putative nucleoside kinase that complexes with SAM68, CIA, NF110/45, and HNRNP-G." Ulke-Lemée A, Trinkle-Mulcahy L, Chaulk S, Bernstein NK, Morrice N, Glover M, Lamond AI, Moorhead GB. Biochim. Biophys. Acta. 2007 Oct;1774 (10):1339-50

(93) "Endofin acts as a Smad anchor for receptor activation in BMP signaling." Shi W, Chang C, Nie S, Xie S, Wan M, Cao X. J. Cell. Sci.. 2007 Apr;120 (Pt 7):1216-24

(94) "PP1 binds Sara and negatively regulates Dpp signaling in Drosophila melanogaster." Bennett D, Alphey L. Nat. Genet.. 2002 Aug;31 (4):419-23